Λ Core Plus also features a new Story mode. The story is a continuation of the storyline from Guilty Gear XX. All characters have a storyline, and the stories cover the major events involving the mystery of Gears, "That Man" and conflict with the Post War Administration Bureau, etc. The mode uses a full branching path Zapping system with a total of 350 scenarios. The game also introduces a new NPC character: the Post War Administration Bureau scientist Crow, who attacks the cast in the Story mode using mindless clones of Justice.

Λ Core Plus also features a new Story mode. The story is a continuation of the storyline from Guilty Gear XX. All characters have a storyline, and the stories cover the major events involving the mystery of Gears, "That Man" and conflict with the Post War Administration Bureau, etc. The mode uses a full branching path Zapping system with a total of 350 scenarios. The game also introduces a new NPC character: the Post War Administration Bureau scientist Crow, who attacks the cast in the Story mode using mindless clones of Justice.